Given a collection of neural networks, can we determine which are parent models and which are child models fine-tuned from the parents?In this work, we strive to answer this questionvia introducing a new task termed as neural phylogeny detection, aimed at identifying the existence and direction of the fine-tuning relationship. Specifically, neural phylogeny detection attempts to identify all parent-child model pairs and determine, within each pair, which model is the parent and which is the child.We present two approaches for neural phylogeny detection: a learning-free method and a learning-based method. First, we propose a metric that leverages the distance from network parameters to a fake initialization to infer fine-tuning directions. By integrating this metric with traditional clustering algorithms, we propose a series of efficient, learning-free neural phylogeny detection methods. Second, we introduce a transformer-based neural phylogeny detector, which significantly enhances detection accuracy through a learning-based manner. Extensive experiments, ranging from shallow fully-connected networks to open-sourced Stable Diffusion and LLaMA models, progressively validate the effectiveness of both methods. The results demonstrate the reliability of both the learning-free and the learning-based approaches across various learning tasks and network architectures, as well as their ability to detect cross-generational phylogeny between ancestor models and their fine-tuned descendants.